Grossly, the gills appear swollen and have a grayish color, caused by the mucus covering the surface. The reduced oxygen uptake may cause an increased rate of opercular movements and fish may gather at the surface of the water, below waterfalls, or near air stones. | Grossly, the gills appear swollen and have a grayish color, caused by the mucus covering the surface. The reduced oxygen uptake may cause an increased rate of opercular movements and fish may gather at the surface of the water, below waterfalls, or near air stones. |