Difference between revisions of "Normal Parturition - Anatomy & Physiology"

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(New page: * The Process by which the conceptus (foetus, placenta and placental membranes) are expelled from the Uterus requires: **...)
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* The Process by which the conceptus (foetus, placenta and placental membranes) are expelled from the [[Female_Reproductive_Tract_-The_Uterus_-_Anatomy_%26_Physiology|Uterus]] requires:
==Stages of Parturition==
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** [[Parturition - Cervical Softening - Anatomy & Physiology|Cervical Softening]]
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** [[Parturition - Coordinated Myometrial Contraction - Anatomy & Physiology|Coordinated Myometrial Contraction]]
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** Voluntary contraction of abdominal muscles
  
*Stage 1: this is the preparatory stage, starting at the onset of regular uterine contractions followed by cervical dilatation and the foetus assuming the correct disposition for passage through the birth canal.
 
*Stage 2: the expulsive stage, characterised by the onset of abdominal contractions which together with uterine contractions lead to foetal expulsion
 
*Stage 3: separation and expulsion of the foetal membranes. In polycotous species second and third stages are often inter-mixed.
 
  
The following table summarises the normal time taken to progress through the stages of parturition in different species.
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* In most species, expulsion of foetal membranes quickly follows expulsion of the foetus. 
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* Expulsion of the foetus requires that the chorionic villi become dislodged from the crypts of the maternal side of the [[Female_Reproductive_Tract_-Placenta_-_Anatomy_%26_Physiology|placenta]].
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** Brought about by powerful vasoconstriction of arteries in the villi.
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*** Vasoconstriction reduces pressure and allows villi to be released from crypts.
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'''Normal Parturition'''
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{| style="width:75%; height:200px" border="1"
 
{| style="width:75%; height:200px" border="1"
  
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| '''Stage 2: Foetal Expulsion'''
 
| '''Stage 2: Foetal Expulsion'''
 
| 12-30 minutes
 
| 12-30 minutes
| 30 minutes - 4 hours
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| 30-60 minutes
 
| 30-120 minutes
 
| 30-120 minutes
 
| 150-180 minutes
 
| 150-180 minutes
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|-
 
|-
 
|}
 
|}
 
 
 
The step in the reproductive process that immediately precedes lactation, uterine involution and return to cyclicity.  It is initiated by the foetus and involves a complex cascade of endocrine events. Parturition is the process by which the conceptus (foetus, placenta and placental membranes) is expelled from the [[Uterus - Anatomy & Physiology|uterus]]; this requires [[Cervical Softening - Anatomy & Physiology|cervical softening]],[[Uterine Contraction - Anatomy & Physiology|coordinated myometrial contractions]] and contraction of abdominal muscles
 
to occur
 
 
== Placental Changes ==
 
 
During the last 5 days of gestation, there are changes in the placenta.
 
 
* Collaginisation of the placentome.
 
* Flattening of maternal crypt epithelium.
 
* Leucocyte migration and increased activity.
 
* Reduction of binucleate cells in the trophectoderm.
 
* Weakening of the acellular protein layer between cotyledonary and caruncular epithelium.
 
 
== Contractions ==
 
 
* Open endometrial crypts
 
* Foetal villi have shrunk due to the escape of blood from the foetal side of the placenta when the umbillical cord ruptures.
 
* [[Uterine Contraction - Anatomy & Physiology|Myometrial contractions]] aid exsanguination of the placenta.
 
* Separation of foetal membranes.
 
* Apex of the allantochorionic sac becomes inverted.
 
* As the sac is 'rolled' down the uterine horns, foetal villi are drawn out of the crypts.
 
* When a large portion becomes detached and inverted, it forms a mass in the maternal pelvis.
 
** Stimulates reflex contractions of abdominal muscles.
 
** Completes expulsion of the allantochorionic sac.
 
* In polytocious species, dehisence and expulsion of foetal membranes are interspersed with births of the young.
 
** Only expulsion of the last afterbirth stimulates abdominal contractions.
 
* The final stage of allantochorionic expulsion lasts 1 hour (mare) - 6 hours (cow).
 
* Domestic animals normally eat the afterbirth.
 
** Not the mare or camelids.
 
 
== Placental Expulsion ==
 
 
In most species, expulsion of foetal membranes quickly follows expulsion of the foetus. 
 
 
* After the birth of the young, regular abdominal contractions largely cease.
 
* Myometrial contractions persist.
 
** Decreased amplitude, but become more frequent and less regular.
 
** Important for dehiscence and expulsion of foetal membranes.
 
* Waves of contractions from uterus to the cervix persist.
 
** Act in a peristaltic fashion in the cow and sow in the reverse direction.
 
 
 
== Parturition Behaviour ==
 
 
=== [[Parturition Behaviour - Mare|Mare]] ===
 
 
===[[Parturition Behaviour - Cow| Cow]] ===
 
 
===[[Parturition Behaviour - Ewe| Ewe]] ===
 
 
===[[Parturition Behaviour - Sow| Sow]] ===
 
 
===[[Parturition Behaviour - Bitch| Bitch]] ===
 
 
===[[Parturition Behaviour - Cat| Cat]] ===
 
 
 
{{OpenPages}}
 
[[Category:Parturition]]
 
[[Category:Bullet Points]]
 

Revision as of 09:11, 15 July 2008


  • In most species, expulsion of foetal membranes quickly follows expulsion of the foetus.
  • Expulsion of the foetus requires that the chorionic villi become dislodged from the crypts of the maternal side of the placenta.
    • Brought about by powerful vasoconstriction of arteries in the villi.
      • Vasoconstriction reduces pressure and allows villi to be released from crypts.


Normal Parturition

Species Mare Cow Ewe Sow Bitch
Stage 1: Contractions and Cervical Dilation 1-4 hours 2-6 hours 2-6 hours 2-12 hours 6-12 hours
Stage 2: Foetal Expulsion 12-30 minutes 30-60 minutes 30-120 minutes 150-180 minutes 6 hours (up to 24 hours with large litters)
Stage 3: Placental Expulsion 1 hour 6-12 hours 5-8 hours 1-4 hours Placenta Exits with Foetus