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**The bile duct and one of the pancreatic ducts discharge into the proximal part of the duodenum by separate ''major duodenal papillae''.
 
**The bile duct and one of the pancreatic ducts discharge into the proximal part of the duodenum by separate ''major duodenal papillae''.
 
**The second pancreatic duct discharges a little more distally on a smaller papilla.
 
**The second pancreatic duct discharges a little more distally on a smaller papilla.
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===Species Differences===
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*Mesoduodenum is relatively short in the horse and ruminant and longer in the carnivore and pig.
      
==Function==
 
==Function==
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*The cranial mesenteric vein drains blood from the duodenum into the portal vein.
 
*The cranial mesenteric vein drains blood from the duodenum into the portal vein.
 
**This blood, carrying the products of digestion, enters the liver.
 
**This blood, carrying the products of digestion, enters the liver.
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==Species Differences==
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===Canine===
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*The duodenum is about 25cm long.
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*The mesoduodenum is relatively long.
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===Horse===
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*Mesoduodenum is relatively short, so the duodenum is closely tethered in a constant position.
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*The descending duodenum is attached to the base of the caecum.
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*The descending duodenum runs caudally beneath the liver to the caudal pole of the right kidney where it has it's caudal flexure to become the ascending duodenum.
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===Ruminant===
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*Mesoduodenum is relatively short.
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*The first part of the descending duodenum rises almost vertically within the ribcage, in close apposition to the liver.
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**It turns level with the tuber coxae.
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*The first part of the duodenum is attached to the liver by the lesser omentum.
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*The descending duodenum also provides attachment for the superficial and deep sheets of the greater omentum.
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===Pig===
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*Small intestine is comparable to the dog.
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*Mesoduodenum is relatively long.
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