Revision as of 11:20, 4 August 2008

Species	Mare	Cow	Ewe	Sow	Bitch
Stage 1: Contractions and Cervical Dilation	1-4 hours	2-6 hours	2-6 hours	2-12 hours	6-12 hours
Stage 2: Foetal Expulsion	12-30 minutes	30-60 minutes	30-120 minutes	150-180 minutes	6 hours (up to 24 hours with large litters)
Stage 3: Placental Expulsion	1 hour	6-12 hours	5-8 hours	1-4 hours	Placenta Exits with Foetus

Parturition is the Process by which the conceptus (foetus, placenta and placental membranes) are expelled from the Uterus requires:

Stage 3: Placental Expulsion

In most species, expulsion of foetal membranes quickly follows expulsion of the foetus.

After the birth of the young, regular abdominal contractions largely cease.
Myometrial contractions persist.
- Decreased amplitude, but become more frequent and less regular.
- Important for dehiscence and expulsion of foetal membranes.
Waves of contractions from uterus to the cervix persist.
- Act in a peristaltic fashion in the cow and sow in the reverse direction.

During the last 5 days of gestation, there are changes in the placenta.

Collaginisation of the placentome.
Flattening of maternal crypt epithelium.
Leucocyte migration and increased activity.
Reduction of binucleate cells in the trophectoderm.
Weakening of the acellular protein layer between cotyledonary and caruncular epithelium.

Open endometrial crypts
Foetal villi have shrunk due to the escape of blood from the foetal side of the placenta when the umbillical cord ruptures.
Myometrial contractions aid exsanguination of the placenta.
Separation of foetal membranes.
Apex of the allantochorionic sac becomes inverted.
As the sac is 'rolled' down the uterine horns, foetal villi are drawn out of the crypts.
When a large portion becomes detached and inverted, it forms a mass in the maternal pelvis.
- Stimulates reflex contractions of abdominal muscles.
- Completes expulsion of the allantochorionic sac.
In polytocious species, dehisence and expulsion of foetal membranes are interspersed with births of the young.
- Only expulsion of the last afterbirth stimulates abdominal contractions.
The final stage of allantochorionic expulsion lasts 1 hour (mare) - 6 hours (cow).
Domestic animals normally eat the afterbirth.
- Not the mare or camelids.

Mammary hypertrophy, waxing of the teats and possible escape of milk from the glands.
Patchy sweating behind the elbows and around the flanks.
- Commences ~4 hours before the birth of the foal and increases as the stage progresses.
Yawns
No indication of pain.
Food is taken readily
Respirations are normal
Pulse is ~60
Body temperature may become slightly subnormal (36.5-37◦C).
Restlessness, wanders aimlessly.
Tail is frequently raised or held to one side.
May swish the tail or slap it against the anus.
Kicking of the abdomen
Towards the end of this stage:
- Crouching
- Straddling of hindlimbs
- Going down on knees or sternum and rising again
- Glancing at flank
This stage terminates with the rupture of the allantochorionic membrane and the escape of allantoic fluid from the vulva.
- No visible straining during this period.

Abrupt onset
Appearance of the amnion or commencement of forceful straining
- Not much delay between them, often coincide.
Soon after straining begins, the mare goes down.
Lies on her side with the limbs extended
- Generally remains in this position until the foal is born.
Transperent blue/white 'water bag' (amnion) visible at the vulva
Quickly followed by the appearance in it of a digit.
Straining at regular intervals.
- Each comprises 3/4 powerful expulsive efforts followed by a period of rest.
- Generally 3 minutes long
One forelimb procedes the other by 7-8cm
- One elbow passes through the pelvic inlet before the other preventing minimal obstruction.
Head is usually in the oblique position, but may be transverse with the cheek lying on the limbs.
Greates and longest effort is with birth of the head.
After expulsion of the foal, the mare may remain on her side exhausted for up to 30 minutes.
Umbillical cord is intact when the foal is born.
- Subsequently ruptures 5-8cm below the belly due to movement of the mare or foal.

@@ Line 111: / Line 111: @@
 === Stage 2 ===
+* Abrupt onset
+* Appearance of the amnion or commencement of forceful straining
+** Not much delay between them, often coincide.
+* Soon after straining begins, the mare goes down.
+* Lies on her side with the limbs extended
+** Generally remains in this position until the foal is born.
+* Transperent blue/white 'water bag' (amnion) visible at the vulva
+* Quickly followed by the appearance in it of a digit.
+* Straining at regular intervals.
+** Each comprises 3/4 powerful expulsive efforts followed by a period of rest.
+** Generally 3 minutes long
+* One forelimb procedes the other by 7-8cm
+** One elbow passes through the pelvic inlet before the other preventing minimal obstruction.
+* Head is usually in the oblique position, but may be transverse with the cheek lying on the limbs.
+* Greates and longest effort is with birth of the head.
+* After expulsion of the foal, the mare may remain on her side exhausted for up to 30 minutes.
+* Umbillical cord is intact when the foal is born.
+** Subsequently ruptures 5-8cm below the belly due to movement of the mare or foal.
 === Stage 3 ===