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**''paneth''  at the base of the crypt. Function unknown. Contain eosinophilic granules.
 
**''paneth''  at the base of the crypt. Function unknown. Contain eosinophilic granules.
 
*Lamina muscularis is smooth muscle.
 
*Lamina muscularis is smooth muscle.
**Contraction of smooth muscle shortens the villus. This helps to pump out absorbed products of digestion. Relaxation of smooth muscle lengthens the villus which increases surface area, facilitating absorption.
  −
*Submucosa is loose connective tissue and it contains blood vessels and nerve fibres.
   
*'''Brunner's glands''' are present in the submucosa of the '''duodenum'''.
 
*'''Brunner's glands''' are present in the submucosa of the '''duodenum'''.
 
**Secrete a protective mucous.
 
**Secrete a protective mucous.
 
**Produce an alkaline secretion which neutralises stomach acid.
 
**Produce an alkaline secretion which neutralises stomach acid.
 
**Open into the crypts in the mucosa above.
 
**Open into the crypts in the mucosa above.
 +
**Contraction of smooth muscle shortens the villus. This helps to pump out absorbed products of digestion. Relaxation of smooth muscle lengthens the villus which increases surface area, facilitating absorption.
 +
*Submucosa is loose connective tissue and it contains blood vessels and nerve fibres.
 
*Tunica muscularis is smooth muscle.
 
*Tunica muscularis is smooth muscle.
 
**It consists of two muscle layers (see regulation & control).
 
**It consists of two muscle layers (see regulation & control).
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