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===Periople===
 
===Periople===
===Coronary Segment===
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===Coronary Dermis Segment===
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The wall of the hoof is created at the coronary dermis and grows in a distal direction from the coronary dermis. The coronary dermis is studded with many papillae which are directed towards the ground in the direction of growth. The epidermis covering these papillae produce horn tubules which are embedded into amorphous inter-tubular horn. This inter-tubular horn is created by the spaces between the papillae within the coronary dermis. The combination of both of these horn types ensures the horn has sufficient strength.
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The pigmentation of the hoof is derived from melanocytes found in the coronary epidermis. Any pigmentation in the hoof will be most pronounced in the outer part of the hoof wall as the deeper layers of the hoof usually contain fewer melanocytes. It is this unpigmented element of the hoof that forms the 'white line' in the sole of hoofs and is particularly important in horses as a landmark for shoeing.
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===Sole Segment===
 
===Sole Segment===
 
===Frog-Bulb Segment===
 
===Frog-Bulb Segment===
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