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==Structures of the Proximal Forelimb and Shoulder==
 
==Structures of the Proximal Forelimb and Shoulder==
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===Radius and Ulna===
 
===Radius and Ulna===
   −
These are complete bones in the ox but are entirely fused. There is a proximal and distal '''interosseous spac'''e which are the only two places where the shafts are seperated. The ulna's proximal end is caudal to the radius and it's distal end forms the '''lateral styloid process''', distal to the radius and articulating with the ulnar carpal bone.
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These are complete bones in the ox but are entirely fused. There is a proximal and distal '''interosseous space''' which are the only two places where the shafts are separated. The ulna's proximal end is caudal to the radius and its distal end forms the '''lateral styloid process''', distal to the radius and articulating with the ulnar carpal bone.
    
==Joints of the Proximal Forelimb==
 
==Joints of the Proximal Forelimb==
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===Carpal bones===
 
===Carpal bones===
   −
Carpal bones comprise two rows:
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The carpal bones comprise two rows:
    
'''Proximally''' (mediolaterally), radial, intermediate, ulnar and accessory bones.
 
'''Proximally''' (mediolaterally), radial, intermediate, ulnar and accessory bones.
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===Metacarpal bones===
 
===Metacarpal bones===
   −
These are covered in detail in the [[Bovine Phalanges - Anatomy & Physiology|bovine phalanges]] section.
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These are covered in detail in the [[Bovine Lower Limb - Anatomy & Physiology|bovine lower limb]] section.
    
==Joints of the Distal Forelimb==
 
==Joints of the Distal Forelimb==
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The carpal joint is a compound joint composed of:
 
The carpal joint is a compound joint composed of:
*The '''antebrachiocarpal joint''' between the radius/ulna and the proximal carpal bones
  −
*The '''middle carpal joint''' between the two rows of carpal bones
  −
*The '''carpometacarpal joint''' between the distal carpal bones and the proximal metacarpals
     −
The joint is a synovial joint, comprised of a common outer fibrous capsule and three inner synovial pouches, one for each joint.  
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1. The '''antebrachiocarpal joint''' between the radius/ulna and the proximal carpal bones.
*'''Collateral ligaments''' extend from the radius to the metacarpal bones on the medial and lateral aspect of the carpus.
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*The carpal canal houses both the deep digital flexor tendon and the deep branch of the superficial digital flexor.
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2. The '''middle carpal joint''' between the two rows of carpal bones.
 +
 
 +
3. The '''carpometacarpal joint''' between the distal carpal bones and the proximal metacarpals.
 +
 
 +
The joint is a synovial joint, consisting of a common outer fibrous capsule and three inner synovial pouches, one for each joint. '''Collateral ligaments''' extend from the radius to the metacarpal bones on the medial and lateral aspect of the carpus. The carpal canal houses both the deep digital flexor tendon, and the deep branch of the superficial digital flexor.
    
==Muscles of the Forelimb==
 
==Muscles of the Forelimb==
 
===Extrinsic Musculature===
 
===Extrinsic Musculature===
   −
These muscle are responsible for joining the forelimb to the trunk, forming a synsarcosis rather than a conventional joint. Collectively, they act to transfer the weight of the body to the forelimbs as well as stabilize the scapula.
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These muscle are responsible for joining the forelimb to the trunk, forming a synsarcosis rather than a conventional joint. Collectively, they act to transfer the weight of the body to the forelimbs as well as stabilize the scapula.
   −
'''''Trapezius''''': Accessory n.
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'''''Trapezius''''':
*Origin: mid-dorsal raphe and supraspinous ligament
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:Innervated by: Accessory n.
*Insertion: spine of the scapula
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:Origin: mid-dorsal raphe and supraspinous ligament.
*Body: two parts, cervical and thoracic separated by aponeurosis
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:Insertion: spine of the scapula.
*Action: raises scapula against the trunk and swings cranially to advance the limb
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:Body: two parts, cervical and thoracic separated by aponeurosis.
 +
:Action: raises scapula against the trunk and swings cranially to advance the limb.
   −
'''''Brachiocephalic m.''''': Accessory n.
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'''''Brachiocephalic m.''''':
*Two parts separated by the clavicle where it exists
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:Innervated by: Accessory n.
*Origin: occipital bone, nuchal ligament, mastoid process
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:The two parts are separated by the clavicle, where it exists.
*Insertion: deltoid tuberosity and fascia of limb
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:Origin: occipital bone, nuchal ligament and the mastoid process.
*Actions:  
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:Insertion: deltoid tuberosity and fascia of the limb.
**advances the limb and extends the shoulder joint when limb is in motion
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:Actions: advances the limb and extends the shoulder joint when limb is in motion. Draws the head and neck ventrally when the limb is fixed.
**draws head and neck ventrally when limb is fixed
     −
'''''Omotransversarius''''': Accessory n.
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'''''Omotransversarius''''':
*Origin: transverse processes of the atlas
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:Innervated by: Accessory n.
*Insertion: acromion and spine of scapula
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:Origin: transverse processes of the atlas.
*Action: advancing the limb
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:Insertion: acromion and spine of scapula.
*Innervation
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:Action: advancing the limb.
   −
'''''Latissimus dorsi''''': local branch of brachial plexus
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'''''Latissimus dorsi''''':
*The '''broadest muscle of the back'''
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:Innervated by: local branch of brachial plexus
*Origin: thoracolumbar fascia  
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:The broadest muscle of the back.
*Insertion: teres tuberosity of the humerus
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:Origin: thoracolumbar fascia.
*Actions: antagonist to the brachiocephalic m.
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:Insertion: teres tuberosity of the humerus.
**cranial fibers strap scapula to the chest
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:Actions: antagonist to the brachiocephalic m. The cranial fibers strap the scapula to the chest. It retracts the free limb and flexes shoulder joint. It draws the trunk forward over the fixed limb.
**retracts free limb and flexes shoulder joint
  −
**draws trunk forward over the fixed limb
     −
'''''Pectoral mm.''''': brachial plexus
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'''''Pectoral mm.''''':
*Two superficial parts, cranial and caudal, these aren't very distinct in the ox
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:Innervated by: brachial plexus
**Origin: cranial sternum
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:Two superficial parts, cranial and caudal, these aren't very distinct in the ox.
**Insertion:
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:Origin: cranial sternum.
***cranial ('''descending'''): crest of the humerus distal to the deltoid tuberosity
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:Insertion:  
***caudal ('''transverse'''): covers elbow joint to insert on the medial fascia of the forearm
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::cranial ('''descending'''), crest of the humerus distal to the deltoid tuberosity.
**Action: adduct the forelimb, assist in protraction and retraction
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::caudal ('''transverse'''): covers elbow joint to insert on the medial fascia of the forearm.
*One deep part ('''pectoralis profundus'''), with cranial and caudal parts
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:Action: adduct the forelimb and assist in protraction and retraction.  
**Origin: ventral sternum and adjacent cartilage
  −
**Insertions:
  −
***cranial (subclavius): supraspinatus m.
  −
***caudal (pectoralis ascendens): lesser tubercle of the humerus
  −
**Actions:
  −
***slinging trunk between forelimbs
  −
***may also retract free limbs
  −
***draw trunk forward when limb is fixed
     −
'''''Serratis ventralis''''': branch of brachial plexus
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'''''Serratis ventralis''''':
*Origin: C4 to 10th rib
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:Innervated by: branch of the brachial plexus.
*Insertion: medial scapula and scapular cartilage
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:Origin: C4 to 10th rib.
*Action: supporting the weight of the trunk
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:Insertion: medial scapula and scapular cartilage.
**reinforced by strong fascia
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:Action: supporting the weight of the trunk. It is reinforced by strong fascia. The cervical portion can retract the limb and the caudal portion can advance the limb.
**cervical portion can retract the limb
  −
**caudal portion can advance the limb
     −
'''''Rhomboids''''': brachial plexus,
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'''''Rhomboids''''':
*Origin: nuchal ligament
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:Innervated by: brachial plexus
*Insertion: dorsal border and adjacent scapula
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:Origin: nuchal ligament.
*Action: retracting the limb, may also raise limb
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:Insertion: dorsal border and adjacent scapula.
 +
:Action: retracting the limb, may also raise the limb.
    
===Intrinsic Musculature===
 
===Intrinsic Musculature===
====Muscles of the Shoulder====
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==Muscles of the Shoulder==
 +
 
 
These muscles are grouped:
 
These muscles are grouped:
*Lateral:'''Supraspinatus ''' and '''Infraspinatus''', Suprascapular n. of the brachial plexus
  −
**Origin: the fossae of the scapula
  −
**Insertion: both tubercles of the humerus
  −
**Action: brace the shoulder
  −
**Clinical significance: bursa between the tendon of the infraspinatus and lateral tubercle of the humerus can be the site of inflammation
  −
*Medial:
  −
**'''Supscapularis''': Subscapular n. from the brachial plexus
  −
***Origin: Deep surface of the scapula
  −
***Insertion: medial tubercle of the humerus
  −
***Action: braces medial shoulder joint, potential adductor
  −
**'''Coracobrachialis''': Musculocutaneous n. of the brachial plexus
  −
***Origin: medial supraglenoid tubercle
  −
***Insertion: proximal shaft of the humerus
  −
***Action: fixator
  −
*Caudal (Flexors): Axillary n. of the brachial plexus
  −
**'''Deltoids'''
  −
***Origin: caudal border and spine of the scapula, acromion
  −
****Two heads of origin
  −
***Insertion: deltoid tuberosity on the humerus
  −
****Action: Flexor of Shoulder
  −
**'''Teres Major'''
  −
***Origin: dorsal part of the caudal scapula
  −
***Insertion: teres tuberosity midway down humerus
  −
**'''Teres Minor'''
  −
*There are no defined extensors of the shoulder.  Those involved (brachiocephalic m., biceps brachii, supraspinatus, and ascending pectorals) have other, more primary roles.
     −
====Muscles of the Elbow====
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===1. Lateral===
''Extensors'': Radial n. from the brachial plexus
  −
*'''Triceps brachii''': Three heads,the medial branch is the most developed
  −
**Long head: caudal margin of the scapula
  −
**Lateral, medial, and accessory heads: shaft of the humerus
  −
**Insertion: olecranon, proteced by tricipital bursa against the bone and subcutaneous bursa against the skin
  −
*'''Tensor fasciae antebrachii'''
  −
**Overlies triceps extending from scapula to olecranon
      +
'''''Supraspinatus''''' and '''''Infraspinatus''''':
 +
:Innervated by: Suprascapular n. of the brachial plexus.
 +
:Origin: the fossae of the scapula.
 +
:Insertion: both tubercles of the humerus.
 +
:Action: brace the shoulder.
 +
:Clinical significance: bursa between the tendon of the infraspinatus and lateral tubercle of the humerus can be the site of inflammation.
   −
''Flexors'': Musculocutaneous n. from the brachial plexus
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===2. Medial===
*'''Biceps brachii'''
  −
**Origin: supraglenoid tubercle of the scapula
  −
**Insertion: medial tuberosity of proximal radius and adjacent ulna
  −
**Runs through the intertubercular groove of the humerus
  −
*'''Brachialis'''
  −
**Origin: proximocaudal humerus
  −
**Insertion: spirals to insert next to biceps
     −
====Muscles of the Carpal and Digital Joints====
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'''''Supscapularis''''':  
''Extensors'': Radial n. from the brachial plexus
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:Innervated by: Subscapular n. from the brachial plexus.
*Craniolateral position on the forearm
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:Origin: Deep surface of the scapula.
*Almost all originate from the lateral epicondyle of the humerus
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:Insertion: medial tubercle of the humerus.
*'''Extensor carpi radialis''': most medial, inserts on 2nd/3rd metacarpal bone
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:Action: braces medial shoulder joint, potential adductor.
*'''Ulnaris lateralis''': most lateral, inserts on accessory carpal bone
  −
*'''Extensor carpi obliquus''': aka '''abductor pollicis longus'''
  −
**Origin: cranial radius
  −
**Insertion: most medial metacarpal bone
  −
*Last two may also serve in medial deviation of the foot
  −
*'''Common Digital Extensor'''
  −
**Insertion: extensor process of the distal phalanx of each digit
  −
*'''Lateral Digital Extensor'''
  −
**Insertion: dorsal proximal phalanges
  −
*'''Medial Digital Extensor'''
  −
**Insertion: middle and distal phalanges
     −
''Flexors'': Median or Ulnar n. of the brachial plexus
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'''''Coracobrachialis''''':
*Caudal position on the forearm
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:Innervated by: Musculocutaneous n. of the brachial plexus.
*Originate from the caudal medial epicondyle of the humerus
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:Origin: medial supraglenoid tubercle.
*'''Flexor carpi radialis''': most medial, inserts on upper 2nd/3rd metacarpal bone
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:Insertion: proximal shaft of the humerus.
*'''Flexor carpi ulnaris''': most lateral, inserts on the accessory carpal bone
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:Action: fixator.
*'''Superficial Digital Flexor'''
  −
**Insertion: palmar surface of middle phalanges
  −
*'''Deep Digital Flexor'''
  −
**Passes through carpal canal before branching and continues to palmar distal phalanges
     −
''Interosseus muscles''
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===3. Caudal (Flexors)===
*This is covered in more detail in the [[Bovine Phalanges - Anatomy & Physiology|bovine phalanges]] section.
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 +
'''''Deltoids''''':
 +
:Innervated by: Axillary n. of the brachial plexus.
 +
:Origin: Two heads of origin; the caudal border and spine of the scapula and the acromion.
 +
:Insertion: deltoid tuberosity on the humerus.
 +
:Action: Flexor of Shoulder.
 +
 
 +
'''''Teres Major''''':
 +
:Innervated by: Axillary n. of the brachial plexus.
 +
:Origin: dorsal part of the caudal scapula.
 +
:Insertion: teres tuberosity midway down humerus.
 +
 
 +
'''''Teres Minor''''':
 +
:Innervated by: Axillary n. of the brachial plexus.
 +
 
 +
 
 +
 
 +
There are no defined extensors of the shoulder. Those involved (brachiocephalic m., biceps brachii, supraspinatus, and ascending pectorals) have other, more primary roles.
 +
 
 +
 
 +
 
 +
==Muscles of the Elbow==
 +
 
 +
===Extensors===
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 +
'''''Triceps brachii''''':
 +
:Innervated by: Radial n. from the brachial plexus
 +
:Has three heads, the medial branch is the most developed. The long head is at the caudal margin of the scapula; The lateral, medial, and accessory heads are at the shaft of the humerus.
 +
:Insertion: olecranon, proteced by tricipital bursa against the bone and subcutaneous bursa against the skin.
 +
 
 +
'''''Tensor fasciae antebrachii''''':
 +
:Innervated by: Radial n. from the brachial plexus.
 +
:Overlies the triceps extending from the scapula to the olecranon.
 +
 
 +
===Flexors===
 +
 
 +
'''''Biceps brachii''''':
 +
:Innervated by:Musculocutaneous n. from the brachial plexus.
 +
:Origin: supraglenoid tubercle of the scapula.
 +
:Insertion: medial tuberosity of proximal radius and adjacent ulna. It runs through the intertubercular groove of the humerus.
 +
 
 +
'''''Brachialis''''':
 +
:Innervated by: Musculocutaneous n. from the brachial plexus.
 +
:Origin: proximocaudal humerus.
 +
:Insertion: spirals to insert next to biceps.
 +
 
 +
==Muscles of the Carpal and Digital Joints==
 +
 
 +
===Extensors===
 +
 
 +
All have innervation from the radial n. from the brachial plexus. They have a craniolateral position on the forearm and almost all of them originate from the lateral epicondyle of the humerus.
 +
 
 +
'''''Extensor carpi radialis'''''
 +
:The most medial, inserts on 2nd/3rd metacarpal bone.
 +
 +
'''''Ulnaris lateralis'''''
 +
:The most lateral, inserts on accessory carpal bone.
 +
 
 +
'''''Extensor carpi obliquus''''' (aka- abductor pollicis longus)
 +
:Origin: cranial radius.
 +
:Insertion: most medial metacarpal bone .
 +
 
 +
'''''Common Digital Extensor'''''
 +
:Insertion: extensor process of the distal phalanx of each digit.
 +
 
 +
'''''Lateral Digital Extensor'''''
 +
:Insertion: dorsal proximal phalanges.
 +
 
 +
'''''Medial Digital Extensor'''''
 +
:Insertion: middle and distal phalanges.
 +
 
 +
===Flexors===
 +
 
 +
All innervated by the median or ulnar n. of the brachial plexus. Have a caudal position on the forearm and originate from the caudal medial epicondyle of the humerus.
 +
 
 +
'''''Flexor carpi radialis'''''
 +
:The most medial, inserts on upper 2nd/3rd metacarpal bone.
 +
 
 +
'''''Flexor carpi ulnaris'''''
 +
:The most lateral, inserts on the accessory carpal bone.
 +
 
 +
'''''Superficial Digital Flexor'''''
 +
:Insertion: palmar surface of middle phalanges.
 +
 
 +
'''''Deep Digital Flexor'''''
 +
:Passes through carpal canal before branching and continues to palmar distal phalanges.
 +
 
 +
'''''Interosseus muscles'''''
 +
:This is covered in more detail in the [[Bovine Lower Limb - Anatomy & Physiology|bovine lower limb]] section.
    
==Vasculature of the Forelimb==
 
==Vasculature of the Forelimb==
*'''[[Arteries of the Forelimb - Anatomy & Physiology|Arteries of the Forelimb]]'''
  −
*'''[[Veins of the Forelimb - Anatomy & Physiology|Veins of the Forelimb]]'''
  −
*'''[[Lymphatics of the Forelimb - Anatomy & Physiology|Lymphatics of the Forelimb]]'''
     −
[[Category:Musculoskeletal System - Anatomy & Physiology]]
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'''[[Arteries of the Forelimb - Anatomy & Physiology|Arteries of the Forelimb]]'''
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'''[[Veins of the Forelimb - Anatomy & Physiology|Veins of the Forelimb]]'''
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'''[[Lymphatics of the Forelimb - Anatomy & Physiology|Lymphatics of the Forelimb]]'''
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