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− | * The Process by which the conceptus (foetus, placenta and placental membranes) are expelled from the [[Female_Reproductive_Tract_-The_Uterus_-_Anatomy_%26_Physiology|Uterus]] requires:
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− | ** [[Parturition - Cervical Softening - Anatomy & Physiology|Cervical Softening]]
| + | ==Stages of Parturition== |
− | ** [[Parturition - Coordinated Myometrial Contraction - Anatomy & Physiology|Coordinated Myometrial Contraction]]
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− | ** Voluntary contraction of abdominal muscles
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− | * In most species, expulsion of foetal membranes quickly follows expulsion of the foetus.
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− | * Expulsion of the foetus requires that the chorionic villi become dislodged from the crypts of the maternal side of the [[Female_Reproductive_Tract_-Placenta_-_Anatomy_%26_Physiology|placenta]].
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− | ** Brought about by powerful vasoconstriction of arteries in the villi.
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− | *** Vasoconstriction reduces pressure and allows villi to be released from crypts.
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| + | *Stage 1: this is the preparatory stage, starting at the onset of regular uterine contractions followed by cervical dilatation and the foetus assuming the correct disposition for passage through the birth canal. |
| + | *Stage 2: the expulsive stage, characterised by the onset of abdominal contractions which together with uterine contractions lead to foetal expulsion |
| + | *Stage 3: separation and expulsion of the foetal membranes. In polycotous species second and third stages are often inter-mixed. |
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| + | The following table summarises the normal time taken to progress through the stages of parturition in different species. |
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| | '''Stage 2: Foetal Expulsion''' | | | '''Stage 2: Foetal Expulsion''' |
| | 12-30 minutes | | | 12-30 minutes |
− | | 30-60 minutes | + | | 30 minutes - 4 hours |
| | 30-120 minutes | | | 30-120 minutes |
| | 150-180 minutes | | | 150-180 minutes |
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| + | The step in the reproductive process that immediately precedes lactation, uterine involution and return to cyclicity. It is initiated by the foetus and involves a complex cascade of endocrine events. Parturition is the process by which the conceptus (foetus, placenta and placental membranes) is expelled from the [[Uterus - Anatomy & Physiology|uterus]]; this requires [[Cervical Softening - Anatomy & Physiology|cervical softening]],[[Uterine Contraction - Anatomy & Physiology|coordinated myometrial contractions]] and contraction of abdominal muscles |
| + | to occur |
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| + | == Placental Changes == |
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| + | During the last 5 days of gestation, there are changes in the placenta. |
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| + | * Collaginisation of the placentome. |
| + | * Flattening of maternal crypt epithelium. |
| + | * Leucocyte migration and increased activity. |
| + | * Reduction of binucleate cells in the trophectoderm. |
| + | * Weakening of the acellular protein layer between cotyledonary and caruncular epithelium. |
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| + | == Contractions == |
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| + | * Open endometrial crypts |
| + | * Foetal villi have shrunk due to the escape of blood from the foetal side of the placenta when the umbillical cord ruptures. |
| + | * [[Uterine Contraction - Anatomy & Physiology|Myometrial contractions]] aid exsanguination of the placenta. |
| + | * Separation of foetal membranes. |
| + | * Apex of the allantochorionic sac becomes inverted. |
| + | * As the sac is 'rolled' down the uterine horns, foetal villi are drawn out of the crypts. |
| + | * When a large portion becomes detached and inverted, it forms a mass in the maternal pelvis. |
| + | ** Stimulates reflex contractions of abdominal muscles. |
| + | ** Completes expulsion of the allantochorionic sac. |
| + | * In polytocious species, dehisence and expulsion of foetal membranes are interspersed with births of the young. |
| + | ** Only expulsion of the last afterbirth stimulates abdominal contractions. |
| + | * The final stage of allantochorionic expulsion lasts 1 hour (mare) - 6 hours (cow). |
| + | * Domestic animals normally eat the afterbirth. |
| + | ** Not the mare or camelids. |
| + | |
| + | == Placental Expulsion == |
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| + | In most species, expulsion of foetal membranes quickly follows expulsion of the foetus. |
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| + | * After the birth of the young, regular abdominal contractions largely cease. |
| + | * Myometrial contractions persist. |
| + | ** Decreased amplitude, but become more frequent and less regular. |
| + | ** Important for dehiscence and expulsion of foetal membranes. |
| + | * Waves of contractions from uterus to the cervix persist. |
| + | ** Act in a peristaltic fashion in the cow and sow in the reverse direction. |
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| + | == Parturition Behaviour == |
| + | |
| + | === [[Parturition Behaviour - Mare|Mare]] === |
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| + | ===[[Parturition Behaviour - Cow| Cow]] === |
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| + | ===[[Parturition Behaviour - Ewe| Ewe]] === |
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| + | ===[[Parturition Behaviour - Sow| Sow]] === |
| + | |
| + | ===[[Parturition Behaviour - Bitch| Bitch]] === |
| + | |
| + | ===[[Parturition Behaviour - Cat| Cat]] === |
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| + | {{OpenPages}} |
| + | [[Category:Parturition]] |
| + | [[Category:Bullet Points]] |