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<big><center>[[Reproductive System|'''BACK TO REPRODUCTIVE SYSTEM''']]</center></big>
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{{OpenPagesTop}}
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==Stages of Parturition==
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<big><center>[[Reproductive_System#Parturition|'''BACK TO PARTURITION''']]</center></big>
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*Stage 1: this is the preparatory stage, starting at the onset of regular uterine contractions followed by cervical dilatation and the foetus assuming the correct disposition for passage through the birth canal.
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*Stage 2: the expulsive stage, characterised by the onset of abdominal contractions which together with uterine contractions lead to foetal expulsion
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*Stage 3: separation and expulsion of the foetal membranes. In polycotous species second and third stages are often inter-mixed.
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The following table summarises the normal time taken to progress through the stages of parturition in different species.
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{| style="width:75%; height:200px" border="1"
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!'''Species'''
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!'''Mare'''
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!'''Cow'''
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!'''Ewe'''
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!'''Sow'''
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!'''Bitch'''
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|-
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| '''Stage 1: Contractions and Cervical Dilation'''
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| 1-4 hours
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| 2-6 hours
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| 2-6 hours
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| 2-12 hours
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| 6-12 hours
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|-
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| '''Stage 2: Foetal Expulsion'''
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| 12-30 minutes
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| 30 minutes - 4 hours
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| 30-120 minutes
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| 150-180 minutes
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| 6 hours (up to 24 hours with large litters)
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|-
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| '''Stage 3: Placental Expulsion'''
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| 1 hour
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| 6-12 hours
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| 5-8 hours
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| 1-4 hours
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| Placenta Exits with Foetus
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|-
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|}
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* The Process by which the conceptus (foetus, placenta and placental membranes) are expelled from the [[Female_Reproductive_Tract_-The_Uterus_-_Anatomy_%26_Physiology|Uterus]] requires:
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** [[Parturition - Cervical Softening - Anatomy & Physiology|Cervical Softening]]
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** [[Parturition - Coordinated Myometrial Contraction - Anatomy & Physiology|Coordinated Myometrial Contraction]]
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** Voluntary contraction of abdominal muscles
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== Placental Expulsion ==
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The step in the reproductive process that immediately precedes lactation, uterine involution and return to cyclicity.  It is initiated by the foetus and involves a complex cascade of endocrine events. Parturition is the process by which the conceptus (foetus, placenta and placental membranes) is expelled from the [[Uterus - Anatomy & Physiology|uterus]]; this requires [[Cervical Softening - Anatomy & Physiology|cervical softening]],[[Uterine Contraction - Anatomy & Physiology|coordinated myometrial contractions]] and contraction of abdominal muscles
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to occur
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In most species, expulsion of foetal membranes quickly follows expulsion of the foetus. 
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== Placental Changes ==
 
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* After the birth of the young, regular abdominal contractions largely cease.
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* Myometrial contractions persist.
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** Decreased amplitude, but become more frequent and less regular.
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** Important for dehiscence and expulsion of foetal membranes.
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* Waves of contractions from uterus to the cervix persist.
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** Act in a peristaltic fashion in the cow and sow in the reverse direction.
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=== Placental Changes ===
      
During the last 5 days of gestation, there are changes in the placenta.
 
During the last 5 days of gestation, there are changes in the placenta.
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* Weakening of the acellular protein layer between cotyledonary and caruncular epithelium.
 
* Weakening of the acellular protein layer between cotyledonary and caruncular epithelium.
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=== Contractions ===
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== Contractions ==
    
* Open endometrial crypts
 
* Open endometrial crypts
 
* Foetal villi have shrunk due to the escape of blood from the foetal side of the placenta when the umbillical cord ruptures.
 
* Foetal villi have shrunk due to the escape of blood from the foetal side of the placenta when the umbillical cord ruptures.
* Myometrial contractions aid exsanguination of the placenta.
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* [[Uterine Contraction - Anatomy & Physiology|Myometrial contractions]] aid exsanguination of the placenta.
 
* Separation of foetal membranes.
 
* Separation of foetal membranes.
 
* Apex of the allantochorionic sac becomes inverted.
 
* Apex of the allantochorionic sac becomes inverted.
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** Not the mare or camelids.
 
** Not the mare or camelids.
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== Placental Expulsion ==
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In most species, expulsion of foetal membranes quickly follows expulsion of the foetus. 
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* After the birth of the young, regular abdominal contractions largely cease.
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* Myometrial contractions persist.
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** Decreased amplitude, but become more frequent and less regular.
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** Important for dehiscence and expulsion of foetal membranes.
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* Waves of contractions from uterus to the cervix persist.
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** Act in a peristaltic fashion in the cow and sow in the reverse direction.
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== Parturition Behaviour ==
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=== [[Parturition Behaviour - Mare|Mare]] ===
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===[[Parturition Behaviour - Cow| Cow]] ===
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===[[Parturition Behaviour - Ewe| Ewe]] ===
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===[[Parturition Behaviour - Sow| Sow]] ===
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===[[Parturition Behaviour - Bitch| Bitch]] ===
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===[[Parturition Behaviour - Cat| Cat]] ===
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{| style="width:75%; height:200px" border="1"
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!'''Species'''
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{{OpenPages}}
!'''Mare'''
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[[Category:Parturition]]
!'''Cow'''
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[[Category:Bullet Points]]
!'''Ewe'''
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!'''Sow'''
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!'''Bitch'''
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|-
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| '''Stage 1: Contractions and Cervical Dilation'''
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| 1-4 hours
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| 2-6 hours
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| 2-6 hours
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| 2-12 hours
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| 6-12 hours
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|-
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| '''Stage 2: Foetal Expulsion'''
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| 12-30 minutes
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| 30-60 minutes
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| 30-120 minutes
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| 150-180 minutes
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| 6 hours (up to 24 hours with large litters)
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|-
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| '''Stage 3: Placental Expulsion'''
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| 1 hour
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| 6-12 hours
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| 5-8 hours
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| 1-4 hours
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| Placenta Exits with Foetus
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|-
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|}
 
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