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* Onset of puberty occurs in January-February in the Northern Hemisphere, where there is increasing day length.
 
* Onset of puberty occurs in January-February in the Northern Hemisphere, where there is increasing day length.
 
* Queens born in February-March do not reach puberty until the following spring.
 
* Queens born in February-March do not reach puberty until the following spring.
* Queens born in sumer/autumn generally display their first oestrus the following January.
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* Queens born in sumer/autumn generally display their first [[Reproductive_Behaviour_-_Oestrus_Behaviour_-_Anatomy_%26_Physiology|oestrus]] the following January.
 
* Before this can have effect, there must be adequate nutrition and growth.
 
* Before this can have effect, there must be adequate nutrition and growth.
  
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