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=== Small strongyles (Cyathostomins) ===
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==== Morphology ====
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'''Gross''':
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*Small worms, <1.5cm long
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*Small, shallow buccal capsule
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'''Microscopic''':
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*Buccal capsule shape
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*Double row of leaf crowns
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*Teeth may be present
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==== Life-cycle ====
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*Infection by ingestion of L3
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*Larvae invade mucosa of large intestine
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*Larvae may develop to L4 without interruption
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*Cyathostomin larvae can arrest at EL3 stage
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*L4 emerge into gut lumen and mature to adult worms
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*Prepatent period 8-12 weeks (depending on species)
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==== Pathogenicity ====
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'''General''':
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*Adult and larval worms are plug feeders, restricting the damage to more superficial mucosa
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'''Cyathostominosis''':
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*Initial infection (L3) → local inflammatory response
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*Developing L4s can be seen as brown flecks in the mucosa
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*They can be present in very large numbers (→ the so-called "pepper-pot lesion")
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*Larval emergence throughout '''summer/autumn''' and plug-feeding of adults → major contributor to the "wormy" horse:
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**Unthriftiness
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**Poor coat
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**Anaemia
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**Diarrhoea)
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*May be tens or hundreds of thousands of adults and millions of mucosal larvae present
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*Emergence of massive numbers of previously arrested larvae in '''late winter/early spring''' → massive inflammatory infiltration → serious disease characterised by severe diarrhoea and/or weight loss (larval or Type 2 cyathostominosis)
       
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