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*Simultaneously, the Cranial air sacs recieve air which was inhaled at the previous inhalation which is drawn from the lungs, this air has lost much of it's oxygen content.
 
*Simultaneously, the Cranial air sacs recieve air which was inhaled at the previous inhalation which is drawn from the lungs, this air has lost much of it's oxygen content.
 
*During Expiration the air sacs are compressed, air from the caudal sac passes through the lungs, while the air in the Cranial Sac leaves via the trachea.
 
*During Expiration the air sacs are compressed, air from the caudal sac passes through the lungs, while the air in the Cranial Sac leaves via the trachea.
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*Thus oxygenated air passes through the lungs on both inspiration and expiration.
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==Avian Gas Exchange==
 
==Avian Gas Exchange==
 
*One '''Primary Bronchi''' from the trachea enters each lung, narrowing as it travels through, and communicates with the '''Abdominal Air Sac'''. This bronchi gives off branches as it travels through the lung, known as '''Secondary Bronchi'''. Each of these gives off a further 400-500 '''Parabronchi''' in the walls of which, gaseous exchange takes place.
 
*One '''Primary Bronchi''' from the trachea enters each lung, narrowing as it travels through, and communicates with the '''Abdominal Air Sac'''. This bronchi gives off branches as it travels through the lung, known as '''Secondary Bronchi'''. Each of these gives off a further 400-500 '''Parabronchi''' in the walls of which, gaseous exchange takes place.
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