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*During Inspiration, the ribs are drawn forwards and the sternum lowered, the Caudal Sacs recieving fresh air.   
 
*During Inspiration, the ribs are drawn forwards and the sternum lowered, the Caudal Sacs recieving fresh air.   
 
*Simultaneously, the Cranial air sacs recieve air which was inhaled at the previous inhalation which is drawn from the lungs, this air has lost much of it's oxygen content.
 
*Simultaneously, the Cranial air sacs recieve air which was inhaled at the previous inhalation which is drawn from the lungs, this air has lost much of it's oxygen content.
*During Expiration the air sacs are compressed, air from the caudal sac passes through the lungs, while the air in the Cranial Sac leaves via the trachea.
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*During Expiration, the sternum is drawn caudal and dorsal, the air sacs are compressed, air from the caudal sac passes through the lungs, while the air in the Cranial Sac leaves via the trachea.
 
*Thus oxygenated air passes through the lungs on both inspiration and expiration.
 
*Thus oxygenated air passes through the lungs on both inspiration and expiration.
  
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